Description
Key Characters:
Growth Form:
Dioecious, weakly erect to sprawling shrubs.
Stems:
Stems often swollen at the nodes, terete (cylindrical), 4–angled or –ridged, 3–6 dm long, few–stemmed from the base, forming tangled clumps, the primary stems branched, weakly ascending to sprawling, internodes 0.8–2.5 cm long, green, glabrous proximally, glabrous to puberulent on the distal stems and inflorescence.
Roots:
Fibrous root system.
Leaves:
Leaves simple.
Opposite, those of axillary stems often appearing fasciculate owing to condensed internodes.
Blades linear or linear–subulate, inconspicuously falcate, weakly v–shaped in cross section, 3–7 (–8.7) cm long, 0.08–0.12 (–0.22) cm wide.
Apex attenuate and often slightly curved.
Base leaf bases often connate around the stem, base slightly tapering.
Surfaces glabrous, but distal ones puberulent on the Upper surfaces near and along the midvein, and on the margins at the base, the hairs 0.05–0.1 mm long, erect, white–opaque; blades, usually reflexed or recurved, becoming reflexed, thin and membranous.
Margins slightly thickened.
Only the midvein evident, the midvein slightly excentric.
Sessile.
Stipules absent.
Flowers:
Flowers in inflorescence terminal, with 15–280 flowers, 6–10 cm long, laterally contracted, lateral branches short, the branches and pedicels ascending to somewhat spreading, glabrous or puberulent with hairs like those of the leaves, spreading to erect; bracts green, filiform to linear–subulate, those of central axis 4.5–6 mm long, those of branches and flowers 1.3–2 mm long.
Flowers unisexual and plants dioecious.
Calyx of staminate flowers with sepals subequal, 2.5–2.65 mm long, narrowly ovate, yellowish green, yellow toward the base, opaque, reflexed and convex at the base, producing a small transverse bulge, the middle part concave, curved upward and broadly and shallowly navicular to nearly flat in the distal ⅓, sometimes irregularly inrolled toward the apex, oriented at 80° to 100° angle to the pedicel, moderately puberulent, the hairs 0.05–0.1 mm long, margins broadly and conspicuously scarious, sometimes ciliate in distal ½, apex obtuse. Pistillate flowers: sepals subequal, 2–2.25 mm long, narrowly ovate, yellowish green, yellow toward the base, opaque, reflexed and convex at the base, producing a small transverse bulge, the middle part concave, curved upward and broadly navicular to nearly flat in the distal ⅓, often irregularly inrolled toward the apex, oriented at 80° to 100° angle to the pedicel, moderately puberulent, the hairs 0.05–0.1 mm long, margins broadly and conspicuously scarious, sometimes ciliate in distal ½, apex obtuse.
Corolla (petals) absent.
Stamens in staminate flowers 10, the filaments dimorphic, 3-5 mm long, the antisepalous whorl 2.8–3.9 mm long, the alternate whorl 2.3–3.5 mm long; anthers ca. 0.3 mm long, pale yellow. Pistillate flowers: Stamens 10 (–11), vestigial, the filaments subequal, 0.55–0.7 mm long; anthers ca. 0.3 mm long, yellowish white, not producing pollen.
Ovary superior (pistillate flowers), 1-celled; ovule placentation free-central; styles 2–3. Staminate flowers: styles 2–3, weakly elongating but non–functional.
Fruit:
Capsules 2–3.5 mm long; narrowly ovoid.
Seeds 0.7–0.8 mm long; reniform; compressed; the surface transversely rugose; perisperm starchy; hard or rarely soft; true endosperm absent.
Ploidy:
2n = 46*–50*
Habitat:
Rare on cliffs and in diverse dry shrubland and mesic forest.
Elevation Range:
60–800 m.